Muscle stem cells undergo extensive clonal drift during tissue growth via meox1-mediated induction of G2 cell-cycle arrest

Date

2017

Authors

Nguyen, P.D.
Gurevich, D.B.
Sonntag, C.
Hersey, L.
Alaei, S.
Nim, H.T.
Siegel, A.
Hall, T.E.
Rossello, F.J.
Boyd, S.E.

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Journal article

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Cell Stem Cell, 2017; 21(1):107-119

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Phong Dang Nguyen, David Baruch Gurevich, Carmen Sonntag, Lucy Hersey, Sara Alaei, Hieu Tri Nim ... al et.

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Abstract

Organ growth requires a careful balance between stem cell self-renewal and lineage commitment to ensure proper tissue expansion. The cellular and molecular mechanisms that mediate this balance are unresolved in most organs, including skeletal muscle. Here we identify a long-lived stem cell pool that mediates growth of the zebrafish myotome. This population exhibits extensive clonal drift, shifting from random deployment of stem cells during development to reliance on a small number of dominant clones to fuel the vast majority of muscle growth. This clonal drift requires Meox1, a homeobox protein that directly inhibits the cell-cycle checkpoint gene ccnb1. Meox1 initiates G<sub>2</sub> cell-cycle arrest within muscle stem cells, and disrupting this G<sub>2</sub> arrest causes premature lineage commitment and the resulting defects in muscle growth. These findings reveal that distinct regulatory mechanisms orchestrate stem cell dynamics during organ growth, beyond the G<sub>0</sub>/G<sub>1</sub> cell-cycle inhibition traditionally associated with maintaining tissue-resident stem cells.

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© 2017 Published by Elsevier Inc.

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